Publications from Wright lab people… [some more pdfs here]

In review

Cornwell WK et al (inc. IJ Wright). Climate and soils together regulate photosynthetic carbon isotope discrimination within C3 plants worldwide.

Gray EF, Wright IJ, Falster DS, Lehmann CER, Bradford M, Cernusak LA. Branch-scale biomass allocation and tissue traits explain variation in stem diameter growth rates of adult trees in a tropical rainforest.

Fernández-Martínez M, Llusià J, Filella I, Niinemets Ü, Arneth A, Wright IJ, Loreto F, Peñuelas J.  Nutrient-rich plants emit a less diverse and intense blend of volatile isoprenoids.

Wang H, Prentice IC, Cornwell WK, Keenan TF, Davis TW, Wright IJ, Evans BJ, Peng C. A universal model for carbon dioxide uptake by plants.

2017 / accepted

Dong N, Prentice IC, Evans B, Caddy-Retalic S, Lowe A, Wright IJ (2017). Leaf nitrogen from first principles: field evidence for adaptive variation with climate. Biogeosciences 14:481-495.

Garnier E, Stahl U, Laporte M-A, Kattge J et al (inc. IJ Wright)(2017). Towards a thesaurus of plant characteristics: an ecological contribution.  Journal of Ecology 105:298-309.

Ghimire B, Riley WJ, Koven CD, Kattge J, Rogers A, Reich PB, Wright IJ (2017). A global trait-based approach to estimate leaf nitrogen functional allocation from observations. Ecological Applications. DOI: 10.1002/eap.1542.

Grootemaat S, Wright IJ, van Bodegom PM, Cornelissen JHC (2017). Scaling up flammability from individual leaves to fuel beds. Oikos  DOI: 10.1111/oik.03886.

Grootemaat S, Wright IJ, van Bodegom PM, Cornelissen JHC, Shaw V (2017). Bark traits, decomposition and flammability of Australian forest trees. Australian Journal of Botany (accepted).

Lu X, Wang Y-P, Wright IJ, Reich PB, Shi Z, Dai Y (2017). Incorporation of plant traits in a land surface model helps explain the global biogeographical distribution of major forest functional types. Global Ecology & Biogeography 26: 304-317.

Onoda Y, Wright IJ, Evans JR, Hikosaka K, Kitajima K, Niinemets Ü, Poorter H, Tosens T, Westoby M (2017). Physiological and structural tradeoffs underlying the leaf economics spectrum. New Phytologist doi: 10.1111/nph.14496.

Scalon MC & Wright IJ (2017). Leaf trait adaptations of xylem-tapping mistletoes and their hosts in sites of contrasting aridity. Plant & Soil  DOI 10.1007/s11104-016-3151-3.

Scalon MC, Wright IJ, Franco AC (2017). To recycle or steal? Nutrient resorption in Australian and Brazilian  mistletoes from three low-phosphorus sites. Oikos 126: 32-39.

Soh WK, Wright IJ, Bacon KL, Lenz TI, Steinthorsdottir M, McElwain JC (2017). Paleo-leaf economics reveal a dramatic shift in ecosystem function at the Triassic-Jurassic boundary. Nature Plants (accepted).

Wang H, Prentice IC, Davis TW, Keenan TF, Wright IJ, Peng C (2017). Photosynthetic responses to altitude: an explanation based on optimality principles. New Phytologist, 213: 976-982.


De Kauwe MG, Lin Y-S, Wright IJ, Medlyn BE, Crous KY, Ellsworth DS, Maire V, Prentice IC, Atkin OK, Rogers A, Niinemets Ü, Serbin SP, Meir P, Uddling J, Togashi HF, Tarvainen L, Weerasinghe LK, Evans BJ, Ishida FY & Domingues TF (2016). A test of the ‘one-point method’ for estimating maximum carboxylation capacity from field-measured, light-saturated photosynthesis. New Phytologist 210:1130-44.

Díaz S, Kattge J, Cornelissen JHC, Wright IJ, Lavorel S et al (2016). The global spectrum of plant form and function. Nature 529:167-171.

Gibert A, Gray EF, Westoby M, Wright IJ, Falster DS (2016). Plant species traits and growth rates: meta-analysis shows correlations change with plant size as predicted. Journal of Ecology 104:1488-1503.

Gleason SM et al (inc. IJ Wright) (2016). Weak tradeoff between xylem safety and xylem-specific hydraulic efficiency across the world’s woody plant species. New Phytologist 209:123-136.

Miatto RC, Wright IJ & Batalha MA (2016). Relationships between soil nutrient status and nutrient-related leaf traits in Brazilian cerrado and seasonal forest communities. Plant & Soil 404:13-33.


Atkin O, Bloomfield K, Reich PB, Tjoelker M et al (inc. IJ Wright)(2015). Global variability in leaf respiration among plant functional types in relation to climate and leaf traits. New Phytologist 206:614-636.

Grootemaat S, Wright IJ, van Bodegom PM, Cornelissen JHC, Cornwell WK (2015). Burn or rot: different roles of leaf traits in fire versus decomposition. Functional Ecology 29:1486-1497. [link]

Maire V, Wright IJ, Prentice IC, Batjes NH, Bhaskar R, van Bodegom PM, Cornwell WK, Ellsworth D, Niinemets Ü, Ordonez A, Reich PB, Santiago LS (2015). Global soil and climate effects on leaf photosynthetic traits and rates. Global Ecology & Biogeography 24:706-717. [link]

Scalon MCWright IJ (2015). A global analysis of water and nitrogen relationships between mistletoes and their hosts: broad-scale tests of old and enduring hypotheses. Functional Ecology 29:1114-1124.  [link]

Schrodt F, Kattge J, Shan H et al (inc. IJ Wright) (2015). BHPMF – a hierarchical Bayesian approach to gap-filling and trait prediction for macroecology and functional biogeography. Global Ecology & Biogeography 24:1510-1521.

Ziemińska K, Westoby M, Wright IJ (2015). Broad anatomical variation within a narrow wood density range — A study of twig wood across 69 Australian angiosperms. PLoS ONE 10:e0124892. [pdf]


Cornwell W, Westoby M, Falster DS et al (inc IJ Wright) (2014). Functional distinctiveness of major plant lineages. Journal of Ecology 102:345-356. [link]

Flores O, Garnier E, Wright IJ et al (2014). An evolutionary perspective on leaf economics in vascular plants: phylogenetic patterns in leaf mass per area. Ecology & Evolution 4:2799-2811. [pdf]

Moles AT, Perkins SE, Laffan SW et al (inc. IJ Wright)(2014). Which is a better predictor of plant traits: temperature or precipitation? Journal of Vegetation Science 25:1167-1180.

Peppe DJ, Lemons CR, Royer DL, Wing SL, Wright IJ, Lusk CH, Rhoden CH (2014). Biomechanical and leaf-climate relationships: a comparison of ferns and seed plants. American Journal of Botany 101:338-347. [link]

Pietsch KA, Ogle K, Cornelissen JHC et al (inc. IJ Wright) (2014). Global relationship of leaf and wood decomposability: the role of trait spectra, phylogeny and decomposability-trait relations across organs. Global Ecology & Biogeography 23:1046-1057.

Prentice IC, Dong N, Gleason SM, Maire V, Wright IJ (2014). Balancing the costs of carbon gain and water transport: testing a new theoretical framework for plant functional ecology. Ecology Letters 17:82-91. [link]

Price CA, Wright IJ, Ackerly DD, Niinemets U, Reich PB, Veneklaas EJ (2014). Are leaf functional traits “invariant” with plant size, and what is “invariance” anyway? Functional Ecology 28:1330-1343.

Richards AE, Wright IJ, Lenz TI, Zanne AE (2014). Sapwood capacitance is greater in woody plant species growing in high compared to low rainfall environments. Functional Ecology 28:734-744.

Zanne AE, Tank DC, Cornwell WK et al (inc IJ Wright) (2014). Into the cold – three keys to radiation of angiosperms into freezing environments. Nature 506:89-92. [link]


Flexas J, Niinemets U, Gallé A, Barbour MM, Centritto M, Diaz-Espejo A, Douthe C, Galmés J, Ribas-Carbo M, Rodriguez PL, Rosselló F, Soolanayakanahally R, Tomas M, Wright IJ, Farquhar GD, Medrano H (2013). Diffusional conductances to CO2 as a target for increasing photosynthesis and photosynthetic water-use efficiency. Photosynthesis Research 117:45–59.

Harrison SP, Morfopoulos C, Dani KGS, Prentice IC, Arneth A, Atwell BJ, Barkley MP, Leishman MR, Loreto F, Medlyn BE, Niinemets Ü, Possell M, Peñuelas J, Wright IJ (2013). Volatile isoprenoid emissions from plastid to planet. New Phytologist 197:49–57.

Kikuzawa K, Onoda Y, Wright IJ, Reich PB (2013). Mechanisms underlying global temperature-related patterns in leaf longevity. Global Ecology & Biogeography 22:982-993.

Maire V, Gross N, Hill D, Martin R, Wirth C, Wright IJ, Soussana J-F (2013). Disentangling coordination among functional traits using an individual-centred model: Impact on plant performance at intra- and inter-specific levels. PLoS One 8:e77372 [pdf].

Mulder C, Ahrestani FS, Bahn M, Bohan DA, Bonkowski M, Griffiths BS, Guicharnaud RA, Kattge J, Krogh PH, Lavorel S, Lewis OT, Mancinelli G, Naeem S, Peñuelas J, Poorter H, Reich PB, Rossi L, Rusch GM, Sardans J, Wright IJ (2013). Connecting the Green and Brown Worlds: Allometric and Stoichiometric Predictability of Above- and Below-Ground Networks. Advances in Ecological Research 49: 69-175.

Perez Harguindeguy N, Diaz S, Garnier E et al (inc. IJ Wright) (2013). New handbook for standardized measurement of plant functional traits worldwide. Australian Journal of Botany 61:167-234. [link]

Verheijen LM, Brovkin V, Aerts R, Bönisch G, Cornelissen JHC, Kattge J, Reich PB, Wright IJ & van Bodegom PM (2013) Impacts of trait variation through observed trait-climate relationships on performance of a representative Earth System model: a conceptual analysis. Biogeosciences 10:5497-5515.

Westoby M, Reich PB, Wright IJ (2013). Understanding ecological variation across species: area-based versus mass-based expression of leaf traits. New Phytologist 199:322-323. [link].

Zieminska K, Gleason SM, Butler DW, Wright IJ, Westoby M. Fibre wall and lumen fractions drive wood density variation across 24 Australian angiosperms. AoB PLANTS 5: plt046. doi:10.1093/aobpla/plt046 [link].


Choat B, Jansen S, Brodribb TJ, Cochard H, Delzon S, Bhaskar R, Bucci SJ, Feild TS, Gleason SM, Hacke UG, Jacobsen AL, Lens F, Maherali H, Martinez-Vilalta J, Mayr S, Mencuccini M, Mitchell PJ, Nardini A, Pittermann J, Pratt RB, Sperry JS, Westoby M, Wright IJ, Zanne AE (2012). Global convergence in the vulnerability of forests to drought. Nature 491:752-755.

Falster DS, Reich PB, Ellsworth DS, Wright IJ, Westoby M, Oleksyn J, Lee TD (2012). Life-time leaf C gain and nutrient use efficiency increases with leaf life-span among 10 Australian woodland species. New Phytologist 193:409-419.

Swenson NG, Enquist BJ, Pither J, Kerkhoff AJ, Boyle B, Weiser MD, Elser JJ, Fagan WF, Forero-Montaña J, Fyllas N, Kraft NJB, Lake JK, Moles AT, Patiño S, Phillips OL, Price CA, Reich PB, Quesada CA, Stegen JC, Valencia R, Wright IJ, Wright SJ, Andelman S, Jørgensen PM, Lacher Jr TE, Monteagudo A, Núñez-Vargas MP, Vasquez-Martínez R, Nolting KM (2012). The biogeography and filtering of woody plant functional diversity in North and South America. Global Ecology & Biogeography 21:798-808.

Tosens T, Niinemets Ü, Westoby M & Wright IJ (2012). Anatomical basis of variation in mesophyll resistance in eastern Australian sclerophylls: news of a long and winding path. Journal of Experimental Botany 63:5105-5119.

Wang YP, Lu XJ, Wright IJ, Dai YJ, Reich PB (2012). Correlations among leaf traits provide a significant constraint on the estimate of global gross primary production. Geophysical Research Letters 39:L19405. doi:10.1029/2012GL053461


Kattge J, Diaz S, Lavorel S, Prentice IC, Leadley P, Bönisch G, Garnier E, Westoby M, Reich PB, Wright IJ and 124 more authors (2011). TRY – a global database of plant functional traits. Global Change Biology 17:2905-2935. [pdf]

Kueffer C, Niinemets Ü, Drenovsky RE, Kattge J, Milberg P, Poorter H, Reich PB, Werner C, Westoby M, Wright IJ (2011). Fame, glory and neglect in meta-analyses. Trends in Ecology & Evolution 26: 493-494. [pdf]

Onoda Y, Westoby M, Adler PB, Choong AMF, Clissold FJ, Cornelissen JHC, Díaz S, Dominy NJ, Elgart A, Enrico L, Fine PVA, Howard JJ, Jalili A, Kitajima K, Kurokawa H, McArthur C, Lucas PW, Markesteijn L, Pérez-Harguindeguy N, Poorter L, Richards L, Santiago LS, Sosinski EE, Van Bael SA, Warton DI, Wright IJ, Wright SJ, Yamashita N (2011). Global patterns of leaf mechanical properties. Ecology Letters 14:301-312. [pdf]

Peppe DJ, Royer DL, Cariglino B, Oliver SY, Newman S, Leight E, Enikolopov G, Fernandez-Burgos M, Herrera F, Adams JM, Correa E, Currano ED, Erickson JM, Hinojosa LF, Hoganson JW, Iglesias A, Jaramillo CA, Johnson KR, Jordan GJ, Kraft NJB, Lovelock EC, Lusk CH, Niinemets Ü, Peñuelas J, Rapson G, Wing SL, Wright IJ (2011). Sensitivity of leaf size and shape to climate: global patterns and paleoclimatic applications. New Phytologist 190:724-739. [pdf]


Ordonez A, Wright IJ, Olff H (2010). Functional differences between native and alien species: a global-scale comparison. Functional Ecology 24:1353-1361. [pdf]

Reich PB, Oleksyn J, Wright IJ, Niklas KJ, Hedin L, Elser JJ (2010). Evidence for a general 2/3-power leaf nitrogen to phosphorus scaling among major plant groups and biomes. Proc. Roy. Soc. Lond. B 277:877-883. [pdf]

Wright SJ, Kitajima K, Kraft NJB, Reich PB, Wright IJ, Bunker DE, Condit R, Dalling JW, Davies SJ, Diaz S, Engelbrecht BMJ, Harms KE, Hubbell SP, Marks CO, Ruiz-Jaen MC, Salvador CM, Zanne AE (2010). Functional traits and the growth-mortality tradeoff in tropical trees. Ecology 91:3664-3674. [pdf]


Cernusak LA, Tcherkez G, Keitel C, Cornwell WK, Santiago LS, Knohl A, Barbour MG, Williams DG, Reich PB, Ellsworth DS, Dawson TE, Griffiths HG, Farquhar GD, Wright IJ & Westoby M (2009) Why are non-photosynthetic tissues generally 13C enriched compared to leaves in C3 plants? Review and synthesis of current hypotheses. Functional Plant Biology, 36:199-213. [pdf]

Craine JM, Elmore AJ, Aidar MPM, Bustamante M, Dawson TE, Hobbie EA, Kahmen A, Mack MC, McLauchlan KK, Michelsen A, Nardoto GB, Pardo LH, Penuelas J,Reich PB, Schuur EAG, Stock WD, Templer PH, Virginia RA,Welker JM & Wright IJ (2009). Global patterns of foliar nitrogen isotopes and their relationships with climate, mycorrhizal fungi, foliar nutrient concentrations, and nitrogen availability. New Phytologist 183:980-992. [pdf]

Hallik L, Niinemets U, Wright IJ (2009). Tolerance to shade and drought in relation to foliage structural and functional traits in temperate woody species. New Phytologist 184:257-274. [pdf]

Moles AT, Wright IJ, Pitman AJ, Murray BG & Westoby M (2009) Is there a latitudinal gradient in seed production? Ecography 32:78-82. [pdf]

Niinemets U, Wright IJ, Evans JR (2009). Leaf internal diffusion conductance in 35 Australian sclerophylls covering a broad range of foliage structural and physiological variation. Journal of Experimental Botany, 60:2433-2449. [pdf]

Ordonez J., Aerts R., van Bodegom P., Witte J-P., Wright IJ, Reich P. (2009) A global study of relationships between leaf traits, climate and soil measures of nutrient fertility. Global Ecology & Biogeography 18:137-149. [pdf]

Poorter H, Niinemets U, Poorter L, Wright IJ, Villar R (2009) Causes and consequences of variation in leaf mass per area (LMA): a meta-analysis. (Tansley review) New Phytologist 182:565-588. [pdf]

Reich PB, Falster DS, Ellsworth DS, Wright IJ, Westoby M, Oleksyn J, Lee TD (2009) Controls on declining carbon balance with leaf age among 10 woody species in Australian woodland: do leaves have zero net carbon balances when they die? New Phytologist, 183:153-166. [pdf]

Reich PB, Oleksyn J, Wright IJ (2009) Leaf phosphorus influences the photosynthesis-nitrogen relation: a crossbiome analysis of 319 species. Oecologia, 160:207-212. [pdf]


Cornwell WK, Cornelissen JHC, Amatangelo K, Dorrepaal E, Eviner VT, Godoy O, Hobbie SE, Hoorens B, Kurokawa H, Perez-Harguindeguy N, Quested HM, Santiago LS, Wardle DA, Wright IJ, Aerts R, Allison S, Van Bodegom PM, Brovkin V, Chatain A, Callaghan T, Diaz S, Garnier E, Gurvich DE, Kazakou E, Klein JA, Read J, Reich PB, Soudzilovskaia NA, Vaieretti MV & Westoby M (2008) Plant species traits are the predominant control on litter decomposition rates within biomes worldwide. Ecology Letters 11: 1065-1071. [pdf]

Poorter L, Paz H, Wright SJ, Ackerly DD, Condit R, Ibarra-Manriquez G, Harms KE, Licona JC, Martinez-Ramos M, Mazer SJ, Muller-Landau HC, Pena-Claros M, Webb CO & Wright IJ (2008) Are functional traits good predictors of demographic rates? Evidence from five Neotropical forests. Ecology 89, 1908-1920. [pdf]

Reich PB, Tjoelker MG, Pregitzer KS, Wright IJ, Oleksyn J & Machado J-L (2008) Scaling of respiration to nitrogen in leaves, stems and roots of higher land plants. Ecology Letters 11, 793-801. [pdf]


Niklas KJ, Cobb ED, Niinemets U, Reich PB, Sellin A, Shipley B & Wright IJ 2007. “Diminishing returns” in the scaling of functional leaf traits across and within species groups. Proc. Nat. Acad. Sci. USA 104, 8891-8896.

Reich PB, Wright IJ & Lusk CH 2007.Predicting leaf physiology from simple plant and climate attributes: A global GLOPNET analysis. Ecological Applications 17, 1982-1988.

Royer DL, Sack L, Wilf P, Lusk CH, Jordan GJ, Niinemets U, Wright IJ,Westoby M, Cariglino B, Coley PD, Cutter AD, Johnson KR, Labandeira CC, Moles AT, Palmer MB & Valladares F 2007.Fossil leaf economics quantified: calibration, Eocene case study, and implications. Paleobiology 33, 574-589.

Wright IJ, Ackerly DD, Bongers F, Harms KE, Ibarra-Manriquez G, Martinez-Ramos M, Mazer SJ, Muller-Landau HC, Paz H, Pitman NCA, Poorter L, Silman MR, Vriesendorp CF, Webb CO, Westoby M & Wright SJ 2007. Relationships among ecologically important dimensions of plant trait variation in seven Neotropical forests. Annals of Botany 99, 1003-1015.


Wright IJ, Falster DS, Pickup M & Westoby M (2006) Cross-species patterns in the coordination between leaf and stem traits, and their implications for plant hydraulics. Physiologia Plantarum 127, 445-456.

Westoby M & Wright IJ (2006) Land-plant ecology on the basis of functional traits. Trends in Ecology & Evolution 21, 261-268.

Lenz TI, Wright IJ & Westoby M (2006) Interrelations among pressure-volume curve traits across species and water availability gradients. Physiologia Plantarum 127, 423-433.

Wright IJ, Leishman MR, Read C & Westoby M (2006) Gradients of light availability and leaf traits with leaf age and canopy position in 28 Australian shrubs and trees. Functional Plant Biology 33, 407-419.

Shipley B, Lechowicz MJ, Wright IJ, & Reich PB (2006) Fundamental tradeoffs generating the worldwide leaf economics spectrum. Ecology 87, 535-541.

Read C, Wright IJ & Westoby M (2006) Scaling up from leaf to canopy-aggregate properties in sclerophyll shrub species. Austral Ecology 31, 310-316.

Warton DI, Wright IJ, Falster DS & Westoby M (2006) Bivariate line fitting methods for allometry. Biological Reviews 81, 259-291.

Wright IJ, Reich PB, Atkin OK, Lusk CH, Tjoelker, MG & Westoby, M (2006) Irradiance, temperature and rainfall influence leaf respiration in woody plants: evidence from comparisons across 20 sites. New Phytologist 169: 309-319.


Wright IJ, Reich PB, Cornelissen JHC, Falster DS, Groom PK, Hikosaka K, Lee W, Lusk CH, Niinemets U, Oleksyn J, Osada N, Poorter H, Warton DI & Westoby M (2005) Modulation of leaf economic traits and trait relationships by climate. Global Ecology & Biogeography 14: 411-421.

Wright IJ, Reich PB, Cornelissen JHC, Falster DS, Garnier E, Hikosaka K, Lamont BB, Lee W, Oleksyn J, Osada N, Poorter H, Villar R, Warton DI & Westoby M (2005) Assessing the generality of global leaf trait relationships. New Phytologist 166: 485-496.

Shipley B, Vile D, Garnier E, Wright IJ & Poorter H (2005) Functional linkages between leaf traits and net photosynthetic rate: reconciling empirical and mechanistic models. Functional Ecology, 19: 602-625.

Vile D, Garnier E, Shipley B, Laurent G, Navas M-L, Roumet C, Lavorel S, Diaz S, Hodgson JG, Lloret F, Midgley GF, Poorter H, Rutherford MC, Wilson PJ & Wright IJ (2005) Specific leaf area and dry matter content estimate thickness in laminar leaves. Annals of Botany 96, 1129-1136.


Wright IJ, Reich PB, Westoby M, Ackerly DD, Baruch Z, Bongers F, Cavender-Bares J, Chapin FS, Cornelissen JHC, Diemer M, Flexas J, Garnier E, Groom PK, Gulias J, Hikosaka K, Lamont BB, Lee T, Lee W, Lusk C, Midgley JJ, Navas M-L, Niinemets U, Oleksyn J, Osada N, Poorter H, Poot P, Prior L, Pyankov VI, Roumet C, Thomas SC, Tjoelker MG, Veneklaas E & Villar R (2004) The world-wide leaf economics spectrum. Nature 428, 821-827.

Wright IJ, Groom PK, Lamont BB, Poot P, Prior LD, Reich PB, Schulze E-D, Veneklaas E & Westoby M (2004) Leaf trait relationships in Australian plant species. Functional Plant Biology 31, 551-558.

Wright IJ & Westoby M (2004) The economics of leaves. Australasian Science 25, 34-38.


Lusk CH, Wright IJ & Reich PB (2003) Photosynthetic differences contribute to competitive advantage of evergreen angiosperm trees over evergreen conifers in productive habitats. New Phytologist 160, 329-336.

Wright IJ, Reich PB & Westoby M (2003) Least-cost input mixtures of water and nitrogen for photosynthesis. American Naturalist 161, 98-111.

Wright IJ & Westoby M (2003) Nutrient concentration, resorption and lifespan: leaf traits of Australian sclerophyll species. Functional Ecology 17, 10-19.

Reich PB, Wright IJ, Cavender-Bares J, Craine JM, Oleksyn J, Westoby M & Walters MB (2003) The evolution of plant functional variation: traits, spectra, and strategies. International Journal of Plant Sciences 164(3 Suppl), S143-S164.

Westoby M & Wright IJ (2003) The spectrum of twig-size variation and its correlates among perennial species in fire-prone sclerophyll vegetation. Oecologia 135, 621-628.


Wright IJ & Westoby M (2002) Leaves at low versus high rainfall: coordination of structure, lifespan and physiology. New Phytologist 155, 403-416.

Wright IJ, Westoby M & Reich PB (2002) Convergence towards higher leaf mass per area has different consequences for leaf lifespan in dry and nutrient-poor habitats. Journal of Ecology 90, 534-543.

Westoby M, Falster DS, Moles AT, Vesk PA & Wright IJ (2002) Plant ecological strategies: some leading dimensions of variation between species. Annual Review of Ecology & Systematics 33, 125-159.


Wright IJ, Reich PB & Westoby M (2001) Strategy-shifts in leaf physiology, structure and nutrient content between species of high and low rainfall, and high and low nutrient habitats. Functional Ecology 15, 423-434.

Wright IJ & Cannon K (2001) Relationships between leaf lifespan and structural defences in a low nutrient, sclerophyll flora. Functional Ecology 15, 351-359.

Wright IJ & Westoby M (2001) Understanding seedling growth relationships through specific leaf area and leaf nitrogen concentration: generalisations across growth forms and growth irradiance. Oecologia 127, 21-29.


Wright IJ & Westoby M (2000) Cross-species relationships between seedling RGR, nitrogen productivity and root versus leaf function in 28 Australian woody species. Functional Ecology 14(1), 97-107.

Wright IJ, Clifford HT, Kidson R, Reed ML, Rice BL & Westoby M (2000) A survey of seed and seedling characters in 1744 Australian dicotyledon species: cross-species trait correlations and correlated trait-shifts within evolutionary lineages. Biological Journal of the Linnean Society 69, 521-547.

Leishman MR, Wright IJ, Moles AT & Westoby M (2000) The evolutionary ecology of seed size. In Seeds – the ecology of regeneration in plant communities, 2nd edition (ed M Fenner), pp 31-57 CAB International, Wallingford, UK.


Wright IJ & Westoby M (1999) Differences in seedling growth behaviour among species: trait correlations and shifts along nutrient compared to rainfall gradients. Journal of Ecology 87(1), 85-97.

Westoby M, Cunningham S, Fonseca C, Overton J & Wright IJ (1998) Phylogeny and variation in light capture area deployed per unit investment in leaves: designs for selecting study species with a view to generalizing. In Inherent Variation in Plant Growth: Physiological Mechanisms and Ecological Consequences (eds H Lambers, H Poorter and MMI Van Vuren), pp 539-566. Backhuys Publishers, Leiden, The Netherlands.

Wright IJ & Ladiges PY (1997) Geographic variation in Eucalyptus diversifolia (Myrtaceae) and the recognition of new subspecies E. diversifolia subsp hesperia and E. diversifolia subsp megacarpa. Australian Systematic Botany 10:651-680.

PUBLICATIONS: Statistics software

Falster DS, Wright IJ 2007/2012. CRU2.0 climate data extraction tool. PrometheusWiki.

Falster DS, Warton DI & Wright IJ (2006). SMATR: Standardised major axis tests and routines – Version 2.0.

Falster DS, Warton DI & Wright IJ (2003). SMATR v1.0.